On Saturday, trains will start late and finish early, and disruption is expected to continue into Sunday morning.
Just 20% of Britain's trains are set to run this Saturday due to strike action and people are being warned to only travel if absolutely necessary.
On Wednesday, Ms Trevelyan told the BBC she wanted unions and train operators to "come together and find something that we can all live with". There will be no direct trains between London and Edinburgh, with passengers told to go via Glasgow. Nor will there be any direct services between London and Norwich.
What makes the RMT strikes so disruptive is that Network Rail's signalling staff across England, Scotland and Wales are involved, so even where train operators are not directly involved in the dispute, only a fraction of usual services are able to run.Tim Shoveller, Network Rail's chief negotiator, said: "Despite our best efforts to compromise and find a breakthrough in talks, rail unions remain intent on continuing and coordinating their strike action.
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