Trains started later than usual on Saturday morning and will finish earlier while some areas will have no services all day.
Events affected by the strike include the Leeds and Reading festivals, Notting Hill Carnival and sports fixtures.
The rail unions are also embroiled in a bitter row over controversial plans to close railway ticket offices, which have angered passenger groups and those representing elderly and disabled passengers. In the letter released on Saturday, RMT general secretary Mick Lynch said: “I believe that both parties are of the view that we need to navigate a way through the dispute and from the RMT’s position that would mean the following suggested stages:
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